靠着母亲的遗言，5年间一直在世界旅行的少女·阳菜。寻找着终结了300年前的大战“废弃战争”的英雄“传说的击落王”的阳菜，在街道上与戴着面具的青年·里希特命运般相遇了。之后，自称击落王的军人出现，将阳菜带到军队……

被自称击落王的士兵欺骗了的阳菜，在不清楚规则的情况下在“星夺战”中败北，自己的数字也被夺走了。里希特瞄准了阳菜母亲的遗物“数字球”，同时也为了快要丢失全部数字的阳菜，里希特展开了对抗！挥舞着刻有白色星纹大刀的里希特的身姿，就如同传说中的击落王？

断崖之街的琳曹长和部下佩雷军曹每天都挥汗于志愿者工作。有一天，他们遇到了和通缉重罪犯里希特长得很像的穿着布丁玩偶服的男人。琳怀疑他是通缉犯，但为了嚎啕大哭着“一点也不受欢迎”的他，琳不得不去和他约会。

琳虽然在追捕者里希特，但还是对住民伸出了援手，看到这一幕的里希特自然而然露出了笑容。但是，原本以为会持续到永远的诙谐逃跑喜剧突然宣告了结束。为了取缔数字球的违法所持者，杰伊尔中尉的铁拳逼近了！

受命取缔违法所持者的中尉杰伊尔，同作为重罪人被通缉的里希特对峙。拥有着里希特两倍以上数字的杰伊尔，能够操纵制造铁的能力，最终捕获了里希特。之后，杰伊尔向里希特提出了在交战中所产生的疑问，试图探寻里希特的真面目。

阳菜和奈奈找到了从杰伊尔率领的皇家军中逃脱后继续一个人旅行的里希特。度过了短暂的平静时光后，他们遇到了在天空中进行飞行实验的女孩珮尔摩。险些坠入谷底的她被里希特救了上来后，向里希特坦白了自己对于飞行的热情。但是，在阿尔西亚进行空中飞行研究是重罪……

里希特一行人来到了集市。里希特和阳菜在街头散步，这时琳出现了。对琳和里希特的亲热，阳菜燃起了对抗之心。奈奈看到两人后大笑，宣布举办阳菜和琳的“争夺里希特料理战斗”。两个人到底做了什么样的料理？

Jail, Lynn and Pele enter the town of Linden and discover that its inhabitants were annihilated by an unknown entity. Jail is confident that Licht must know something about this.

The Demon of the Abyss shows itself. Nobody knows what it is or how to defeat it, except for Licht, who takes command.

Licht unleashes his true power, becoming an overwhelming killing machine. However, it seems like his unmatched strengths come at a high cost.

Licht and Mizuka have returned to their normal selves. While the ladies in the group suffer the consequences of Licht and Mizuka’s unresolved Star Stake, Jail receives an order to report to the supreme commander of the military.

Nana sends Jail, Lynn, Pele and Hina 300 years back in time. They arrive at an unknown institution with technology far superior to the world they are accustomed to, and meet Rihito, a young man Hina is convinced is Licht himself.

The time travelers settle into their new role as students in the military academy. However, when Schmerman reveals the "rules" of the school, it becomes apparent that there’s more to this place than fun times and friendship.

A voyeur is regularly sneaking a look at the girls’ bath, and the students are tasked with catching the culprit. Meanwhile, Jail attempts to find out more about this timeline and figure out why Nana sent them here.

Jail discovers more secrets about this world through Nana, but is intercepted by Alan during his escape. Meanwhile, the students of Class A come closer to realizing their dream of becoming a military force that does not take human lives.

Rihito decides to receive the operation to turn him into an Ace. This is the decisive moment in history that the time travelers wanted to change, but knowing what they know now, they cannot agree on what is best for everyone.

Jail and the squad are successful. With his newfound overwhelming powers, Rihito should be able to defeat the enemy with ease. However, the enemy, a group of trained and determined soldiers, will not go down without resistance.

The time travelers return to the future, and learn the true history of their world. Though they were unable to change the past, they are determined to correct the present and shape the future.

Nana throws a party for the weary time travelers after their long journey through history. One by one, they recover from the gloomy atmosphere brought on by what they had witnessed.

Licht takes off on his own and faces off against Doan, whose power completely overwhelms him. The others chase after him, while Jail visits his adopted father to seek closure.

Licht is dead. Doan determines that the others are also attempting to take down Alcia, so he decides to kill them all as well.

Like he did in the past, Licht returns to life from what apparent death, and returns to save the others from Doan. However, Mizuka joins Doan. Licht must take both Aces on at once.

With Mizuka subdued, the only opposition facing Licht is Doan. Doan traps Licht in a prison of gravity, from which there is no escape. Hina, however, still believes in Licht.