When Leslie decides to marry two penguins to promote the local zoo, she inadvertently causes an uproar when both penguins turn out to be male. Meanwhile, Mark puts Ann in an awkward situation when he asks her to go see a movie.

Leslie wants to bury a time capsule that encapsulates life in Pawnee, but an odd suggestion from a local man causes unforeseen consequences. Andy seeks help from Chris about how to win back April.

Jerry’s contribution to Leslie’s community art show causes a stir. Ben moves in with April and Andy and attempts to teach them how to be adults. Meanwhile, Leslie is frustrated by Chris and his interoffice dating rules.

An STD outbreak amongst Pawnee seniors prompts Leslie to conduct a sex education class. Meanwhile, Ron helps Tom adjust to life without technology after his gadgets are taken away, and Ben and April meet Congressman Murray for the first time.

Leslie celebrates her first year as a city councilwoman. Retired Special Agent Burt Macklin is recalled to action. April gets some life-changing news. Tom debates a big business opportunity.

Leslie and Ben come under fire for participating in an old Pawnee tradition. April and Ron go on a scavenger hunt. Donna and Garry stroll down memory lane.