Desna and the girls struggle to keep hope alive while still working under the thumb of the new Russian regime, spearheaded by Riva. Desna remains in the dark about her gangster/gyno boyfriend, Ruval. Virginia tells Dean that she's pregnant.

Desna, now pinned under the stiletto-tipped nail of Riva's sister, Zlata, must come up with an idea to generate more customers for the clinic. Quiet Ann helps Dean and Virginia get to a very important appointment.

Desna's relationship with Ruval deepens when he introduces her to his mother, Matilde. Polly grows concerned when Marnie continues to pull away from her. Jenn, still depressed by Bryce's recent indiscretion, falls off the wagon.

Desna, in an attempt to cozy up to Matilde, Ruval's mother, puts together a spa day for her and a couple of young girls from his charity. Quiet Ann gets a surprise visit from her brother, who convinces her to attend a family dinner.

Desna continues to grow closer to Zlata, despite the crew's increasing concern as her relationship with Dr. Ruval intensifies. Meanwhile, Dean gets an opportunity to showcase his talents and Uncle Daddy gets his mojo back.

Faced with a dilemma, Desna and Dean take a road trip to seek guidance from a childhood mentor. Back in Palmetto, Jenn must lean on Polly and Quiet Ann for help with a personal crisis. Virginia struggles to run the salon on her own.

When Desna with the help of Uncle Daddy and her crew plots revenge on those who betrayed her, she realizes that vengeance comes at a high cost.

Desna is offered a deal with the Russian devil, aka Zlata, that's more intriguing than nail art on acrylics. Polly and Quiet Ann get dealt some hard-ass betrayal from their baes while Jenn's affair comes back to haunt her.

Polly gets an unexpected visit from her long-lost sister that sends her spinning out of control. Uncle Daddy and Roller join forces with their Dixie Mafia brethren and make a plan to take out the Russian and Haitian mafias for good.