Jane and Michael's wedding plans are side tracked when the Villanueva house is ruined by a flood. Jane is looking to have a small wedding, but Rogelio wants nothing more than to throw Jane a big blow out wedding so he can invite all his celebrity friends. Jane and Michael are also looking for the perfect home to move into and when they find a place 45 minutes away, Rafael has an issue being that far from Mateo. Petra is still having trouble adjusting to motherhood.

Jane takes on additional shifts at the Marbella to make some extra money, but it starts cutting into her time with Mateo. Jane and Rafael agree that they need a formal custody agreement and decide to use a mediator to help make it more amicable. Xo takes an acting role on Rogelio's telenovela and they are both surprised with her performance. Michael gets called into his boss' office after a newspaper runs a story on him and Sin Rostro. Meanwhile, Petra confronts Rafael when she learns that the twins' trust fund is different from Jane's arrangement.

Jane has made big plans to celebrate Mateo’s first birthday, but things start to fall apart when she learns there might be an issue with her wedding venue. Rogelio faces a challenge on set that could jeopardize Jane’s big day. Jane is excited when Alba gives her a lead on a book publisher, but soon begins having doubts about her writing. Jane and Petra try to help Rafael figure out his troubles, but they have differing opinions on how to handle the situation. Meanwhile, Michael is convinced that Derek is really in cahoots with Mutter and has a plan to prove it.

Jane continues to work on her thesis and she decides to add Alba’s estranged sister to the narrative, much to Alba’s dismay. In order for Rogelio to have a chance at being an American crossover star he decides to bring the Passions of Santos to The CW in hopes they will pick it up. Xo thinks about abandoning her dream of being a singer for something more realistic. Meanwhile, Luisa is forced to make a choice between her family or Rose.

When Jane’s cousin unexpectedly shows up at her door, Jane questions her own less than spontaneous life. Xo finds the perfect location for her dance studio, but it happens to be near her ex-boyfriend, Bruce. Rogelio realizes his feelings for Xo, but struggles with telling her. Petra is served with a lawsuit and she decides to ask Rafael for help. Meanwhile, Michael is ready to get to work just as they get a break in the Mutter case.

Jane is finally a published author but she’s reluctant to talk about her past to help sell her book. Rogelio is intimidated by his new younger male co-star and begins to wonder if the newbie is trying to sabotage him. Luisa and her girlfriend Eileen are back in town to visit Rafael, but Rafael isn’t taking any chances when it comes to safety. Meanwhile, Petra must convince Anezka to come back to Miami to answer some questions the police have for her.

Jane is ready for some casual dating and is pleasantly surprised when her date actually helps her fix things with Lina. Petra follows a lead with the help of Chuck that could help her sister’s case. Rafael takes Jane’s advice and tries to come clean with Petra. Rogelio is angry with Xo when she blows him off, but is shocked when he learns the reason why. Meanwhile, Alba is appalled by a comment a customer makes in the gift shop and decides to take a political stance.

Jane has to learn how to handle her new love life, especially when the guy she is dating introduces her to his grandmother. Petra learns some interesting information about Chuck. Xo and Rogelio are eager to share their news with Jane, but want to try to take her feelings into consideration.

Meanwhile, when Mateo starts asking questions, Jane and Rafael are forced to sit down with him and explain how their family came to be.

When Jane is reunited with her first love, Adam, she is shocked but happy to see him, however Xo and Alba are less thrilled that he is back. After Rafael loses ownership of the hotel to his sister, he finds himself at Jane's house, broke and homeless. Jane and Rafael learn that Petra has been found and they rush to her aide, but Anezka is nowhere to be found. Meanwhile, Xo and Rogelio wedded bliss is interrupted when they confront the realization that Darci is having Rogelio's baby and she wants full custody.

Jane and Adam's relationship seems to be moving at hyper-speed, and Jane thinks that it might be time for him to meet Mateo, but Rafael is not happy with this idea. As the arrival of baby De LaVega Factor approaches, Rogelio's jealously of Darci's new boyfriend increases as well. Trying to be supportive, Xo encourages Rogelio to step up and be the father he was meant to be. Meanwhile, Rafael is determined to get his hotel and life back and will do whatever he has to do to make it happen.

When Adam reveals his dating past to Jane, the information puts a strain on their relationship. Lina seeks Jane's advice about whether she should get married and begs Jane to give her honest opinion about her fiancé. Petra hears some important information that could help Rafael get the hotel back, but Rafael is more concerned about his sister's well-being. Meanwhile, Xo asks Rogelio to consider having a medical procedure done, but Rogelio is very hesitant.

Jane's book is finally being released but her plans to have a party at the local bookshop where she was inspired to be a writer hit a snag. Always wanting to help Jane, Rogelio devises a plan to save the day, but one of his archrivals threatens to ruin the day. Rafael finds a job, but it is not exactly what he was hoping for. Meanwhile, Petra has come up with an interesting strategy to get the Marbella back, but Rafael wants nothing to do with it.

Jane and Rafael find themselves in a not ideal living situation in order to save money. When Rafael sees Alba try to punish Mateo, an unexpected rift is formed. Xo tries to stay positive during treatment and even makes a new friend, but things get a little too real for her and Rogelio. River and Rogelio once again are at odds over their ideas for the show, but an unlikely person helps River see the show in a different light. Meanwhile, Petra feels unsupported by JR when she questions what really happened the night Anezka died.