Jane and Rafael are still at odds with each other as they are about to celebrate Mateo's first Christmas. Jane discovers that the scholarship she thought she had been awarded isn't quite that and now needs to find a way to pay for grad school. Rogelio has employed an intern and is having trouble trusting him. Petra is struggling with the latest dilemma her mother has gotten them into. Meanwhile, Michael is trying to find Luisa's mother who he now suspects is not dead and may be involved with Sin Rostro.

Jane's latest parenting obstacle is sleep training Mateo, a struggle that has the Villanueva woman divided on what is the best method. At the same time, she is beginning her new job as a TA and is faced with a class full of basketball players looking for an easy A. Petra confesses all to Rafael and is determined to put her mother away for good. Rogelio's mother Liliana comes for a visit and reveals a 40 year old family secret. Meanwhile, Michael discovers the truth about Luisa's mother and new information about Rafael's mother.

With Jane and Xo still dealing with the fallout from Jane's bachelorette party, Jane is now left with finding someone to watch Mateo. When Michael volunteers to help everything is perfect until Mateo does something for the first time in front of Rafael, upsetting him. Rogelio gets in trouble with the network executives when he uses the crew for his own use, so he turns to his favorite writer Dina to help fix the problem. Meanwhile, someone is causing problems for Petra at the Marbella and in her personal life. 

Jane takes on additional shifts at the Marbella to make some extra money, but it starts cutting into her time with Mateo. Jane and Rafael agree that they need a formal custody agreement and decide to use a mediator to help make it more amicable. Xo takes an acting role on Rogelio's telenovela and they are both surprised with her performance. Michael gets called into his boss' office after a newspaper runs a story on him and Sin Rostro. Meanwhile, Petra confronts Rafael when she learns that the twins' trust fund is different from Jane's arrangement.

The Villanueva women have a tradition for Mother's Day and Jane is excited to get to participate in her first one as a mom. However, their plans are thwarted when Petra invites everyone to a brunch instead. Jane's TA position is put into jeopardy when a mysterious ad appears in the college newspaper. Rogelio is hurt when Dina doesn't invite him to her birthday party, so he asks Jane to help in an unusual way. Rafael finds himself in big trouble thanks to his half-brother. Meanwhile, Xo finds herself in a compromising position.

Jane and Michael’s housewarming party in their new home hits a snag when they discover they are being evicted for not paying their rent. Rogelio is still trying to break into American television, but needs some unusual help from Rafael. Xo makes a new career choice, but Alba and Jane aren’t convinced this is the one for her. Meanwhile, “Petra” shocks Rafael with what she wants to do with her shares of the hotel.

When Jane’s cousin unexpectedly shows up at her door, Jane questions her own less than spontaneous life. Xo finds the perfect location for her dance studio, but it happens to be near her ex-boyfriend, Bruce. Rogelio realizes his feelings for Xo, but struggles with telling her. Petra is served with a lawsuit and she decides to ask Rafael for help. Meanwhile, Michael is ready to get to work just as they get a break in the Mutter case.

Jane and Fabian are not on the same page when it comes to their relationship, which makes it hard for Jane to ask him for a favor. Petra and Rafael both turn to Jane for advice, but Jane is determined not to get involved. Meanwhile, Jane volunteers to help Xo and Rogelio, but might have bitten off more than she can chew.

Jane and Adam's relationship seems to be moving at hyper-speed, and Jane thinks that it might be time for him to meet Mateo, but Rafael is not happy with this idea. As the arrival of baby De LaVega Factor approaches, Rogelio's jealously of Darci's new boyfriend increases as well. Trying to be supportive, Xo encourages Rogelio to step up and be the father he was meant to be. Meanwhile, Rafael is determined to get his hotel and life back and will do whatever he has to do to make it happen.

When Adam reveals his dating past to Jane, the information puts a strain on their relationship. Lina seeks Jane's advice about whether she should get married and begs Jane to give her honest opinion about her fiancé. Petra hears some important information that could help Rafael get the hotel back, but Rafael is more concerned about his sister's well-being. Meanwhile, Xo asks Rogelio to consider having a medical procedure done, but Rogelio is very hesitant.