Michelle Webster, the rebellious teenage daughter of a friend of MacLeod's, drives away from her parents' house in a rage and right over a cliff. Trauma surgeon Anne Lindsey tries her best to save Michelle, but it's too late. MacLeod rushes to the hospital to comfort his grieving friends -- and sneak their newly Immortal daughter out of the morgue. He tries to train her in the arts of Immortality, but Michelle just wants to have fun. She meets Immortal Axel Whittaker who promises her all the fun and adventure she could imagine if she stays with him. In flashback, we see that Axel uses beautiful new Immortal women as bait to trap other Immortals and take their heads -- MacLeod barely escaped with his in 1896 Boston. Axel uses Michelle to lure MacLeod to his yacht, where they continue the battle they started a hundred years before. MacLeod defeats Axel. Michelle, witnessing the fearsome power of the Quickening, agrees to be trained as an Immortal under the protection of Amanda.

Father Bernard has a secret he thought he'd buried 50 years ago at the bottom of the Seine. When Ernest Daimler, the Nazi Major who Bernard killed as a young boy and threw in the river, appears at his church looking not a day older than the day he died, Father Bernard realizes he's like MacLeod. As a child, Bernard watched MacLeod die and revive during a mission for the French Resistance and MacLeod swore him to secrecy. Bernard goes to MacLeod for help. Meanwhile, Anne tells MacLeod that she is pregnant -- by an old friend she sought for comfort after MacLeod's ""death."" MacLeod tries to adjust to the concept of being a father. When Daimler kills Father Bernard and comes after Anne, MacLeod kills Daimler and Anne witnesses what being part of MacLeod's life really means. Unable to deal with the part of herself that wanted to see Daimler die, Anne leaves MacLeod.

Amanda accidentally helps Kalas escape from prison by trying to do MacLeod a favor and kill Kalas for him. After an attempt on Maurice's life thwarted by MacLeod, Kalas kidnaps Amanda to use as bait against MacLeod, but Amanda manages to escape. Meanwhile, Christine Salzer, the widow of a Watcher killed by Kalas (in ""Methos"") decides to get her revenge on Immortals and Watchers alike by exposing their secret to the media. Dawson and Methos team up to try and talk Christine out of it, but she's determined to take a computer disk with the identities of all known Immortals and Watchers on it to a newspaper publisher. Dawson, desperate, tries to kill her outside the newspaper building, but is stopped by MacLeod and Methos. Christine enters the building and the Immortals and the Watcher know their lives are about to change forever.

The past comes back to haunt Richie when he spies Mark Roszca, the street punk who killed Tessa. Richie is determined to go after Roszca and revenge Tessa's death. Meanwhile, MacLeod is distracted by the return of an old Immortal enemy, Kanis -- a master of hounds who uses his pack of dogs to track and exhaust his prey. This time his prey is Duncan MacLeod.

MacLeod's old friend, Kit O'Brady, comes into town in search of a good race horse and a change of luck. MacLeod knew Kit back in Gold Rush San Francisco, when Kit ran the Double Eagle Saloon. Kit lost the Double Eagle to Amanda in a poker game and has blamed her ever since for his string of bad luck -- and he's vowed to kill her. When Amanda chooses to breeze back into MacLeod's life while Kit is there, MacLeod is hard pressed to keep them apart and to keep them from killing each other.