The department is still reeling with the recent departure of Deputy Chief Brenda Lee Johnson, creating a tough transition for the newly appointed head of Major Crimes, Captain Sharon Raydor. But there is little time for adjustment as the department rushes to reach a plea bargain when a string of grocery store robberies turns fatal. After proving to be a valuable source for the case, undercover police detective and military veteran Detective Amy Sykes is transferred to Major Crimes. Meanwhile, Raydor takes a big step to protect teenager Rusty Beck after he runs away from his foster family.

Growing pains within Major Crimes become apparent as the division works to obtain justice in a brutal homicide that turns out to be more complicated than expected. Raydor struggles to gain the respect of her colleagues while also trying to supervise Rusty 24/7. The changes within the department may become more than Provenza can accept.

The facts become blurry in a case involving a fatal car crash at a nightclub. Meanwhile, Rusty can finally look forward to his mother's return.

The unit joins forces with the FBI when a homicide case spins around the Israeli mob and an "intuitive life strategist." Elsewhere, Provenza displays his vulnerable side, and Rusty's first school day takes an unexpected turn.

When a kidnapping turns deadly, the squad scrambles to crack the case and turns to a district attorney for help. Meanwhile, DNA evidence shows that Rusty has connections to the justice system.

The aftermath of a deadly shooting on gang turf finds Raydor making a daring maneuver that could threaten her career and jeopardize Det. Sykes' life. Elsewhere, Rusty makes it quite apparent that he has no interest in meeting his biological father.

When a murder conviction is overturned after eight years, Tao is forced to question his past actions. And Rusty struggles to be himself during his first outing with his father, Daniel Dunn.

A man from Las Vegas is found dead after an apparent overdose, but the facts become unclear as gritty details begin to unfold. Meanwhile, a series of strange events leave Rusty and Raydor wondering if Dunn (Ian Bohen) can be trusted.

Season Finale. When a peaceful ceremony turns deadly, Major Crimes and the FBI team up to track down a dangerous sniper. And Raydor negotiates a deal that even Provenza can get behind.