The department is still reeling with the recent departure of Deputy Chief Brenda Lee Johnson, creating a tough transition for the newly appointed head of Major Crimes, Captain Sharon Raydor. But there is little time for adjustment as the department rushes to reach a plea bargain when a string of grocery store robberies turns fatal. After proving to be a valuable source for the case, undercover police detective and military veteran Detective Amy Sykes is transferred to Major Crimes. Meanwhile, Raydor takes a big step to protect teenager Rusty Beck after he runs away from his foster family.

Growing pains within Major Crimes become apparent as the division works to obtain justice in a brutal homicide that turns out to be more complicated than expected. Raydor struggles to gain the respect of her colleagues while also trying to supervise Rusty 24/7. The changes within the department may become more than Provenza can accept.

The facts become blurry in a case involving a fatal car crash at a nightclub. Meanwhile, Rusty can finally look forward to his mother's return.

When a kidnapping turns deadly, the squad scrambles to crack the case and turns to a district attorney for help. Meanwhile, DNA evidence shows that Rusty has connections to the justice system.

The aftermath of a deadly shooting on gang turf finds Raydor making a daring maneuver that could threaten her career and jeopardize Det. Sykes' life. Elsewhere, Rusty makes it quite apparent that he has no interest in meeting his biological father.

When an activist to end human trafficking is murdered, the investigation leads to a possible eyewitness, a young prostitute wanting to get out of the business, and the pimp she works for becomes their primary suspect. Meanwhile, child services arrange a meeting between Rusty and his father, against Rusty's wishes.

When a murder conviction is overturned after eight years, Tao is forced to question his past actions. And Rusty struggles to be himself during his first outing with his father, Daniel Dunn.

A man from Las Vegas is found dead after an apparent overdose, but the facts become unclear as gritty details begin to unfold. Meanwhile, a series of strange events leave Rusty and Raydor wondering if Dunn (Ian Bohen) can be trusted.

Season Finale. When a peaceful ceremony turns deadly, Major Crimes and the FBI team up to track down a dangerous sniper. And Raydor negotiates a deal that even Provenza can get behind.