Some of the team have to assume undercover identities in Moscow so that they can locate their suspects.

While they’re in Russia, Callen, Sam and Anna craft a plan to break Anna’s father and his CIA agent companion out of prison. Also, Special Agent G. Callen finally learns what the “G” in his name stands for, and in Los Angeles, Kensi and Deeks take Eric into the field on an assignment.

After Callen’s father, Garrison is found in an NCIS patient’s hospital room during a radiation poisoning case, Callen must take him to the boatshed for interrogation. Also, the team discusses their Thanksgiving plans.

After Anna Kolcheck escapes from prison with her cellmate, Callen and the NCIS team assist with the investigation and search for her whereabouts.

The NCIS team travels to Cuba for an unsanctioned mission after Callen gets a lead on Anna Kolcheck's (Bar Paly) whereabouts.

Callen and Sam work with Navy Captain Harmon "Harm" Rabb, Jr. (David James Elliott) to locate a group of ISIS sympathizers aboard the USS Allegiance. Also, Hetty enlists her friend, Lieutenant Colonel Sarah "Mac". Mackenzie (Catherine Bell), to help obtain information from a Russian diplomat.

Anna Kolcheck returns to warn Callen that he is in danger and he must now work with an archenemy to stop an underground trafficking ring.