When Echo, Sierra and Victor awaken in the Dollhouse with most of their original personalities and memories intact, Echo leads the charge to free the Actives from their apparent captivity and escape the Dollhouse forever. Meanwhile, Ballard discovers the Dollhouse is closer than he thinks.

Echo is imprinted with the personality of a mother with a newborn baby. Complications arise when, thanks to Topher's programming, Echo takes too strongly to motherhood. Perrin ramps up his investigation into Rossum and DeWitt pays November a visit.

When Senator Perrin makes his stand against the Rossum Corporation, Echo attempts to stop his expose. Adelle, and Topher begin to see what they are up against in the rival of Washington DC.

Echo and Bennett have a shocking meeting as Adelle goes head-to-head with the ruthless head of the DC Dollhouse. Topher sees double when he must involve Victor in his espionage. Perrin finds a surprising witness to testify against the Rossum Corporation, but find he may be living in a house of cards.

Topher races to reconstruct Echo's original personality and memories when Adelle realizes they are the key to averting a devastating future. Meanwhile, Echo and Bennett's shared past and the identity of The Rossum Corporation's mysterious leader are revealed.

Sacrifices are made when Echo leads her crew to Arizona to dismantle the Rossum Corporation's mainframe.