Upon Mr. Dashwood's death the entirety of his estate passes, by law, to his son John from his first marriage, leaving his widow and daughters entirely dependent on the latter's reluctant generosity. Now mere guests in their own home, Mr. Dashwood's daughters meet visiting relative-in-law Edward Ferrars. He and eldest daughter Elinor Dashwood are drawn to each other, but Elinor learns Edward's inheritance is conditioned on his marrying well—and knows her relative poverty disqualifies her.

In London, Willoughby acknowledges no connection to Marianne and marries wealthy heiress Grey. Elinor must hide her heartbreak when she learns Edward Ferrars is engaged to Miss Lucy Steel; when his mother learns of the engagement she disinherits him. Fortune smiles unexpectedly on both Dashwood sisters.