When Rumpelstiltskin destroys the Magic Mirror and escapes to the modern world, the four princesses of "Once Upon a Time"-Cinderella, Sleeping Beauty, Snow White, and Rapunzel-are sucked through the portal too. Well-trained and endowed with magical powers, the four women must fight Rumpelstiltskin and his army of thralls before he enslaves everyone on Earth.

An alliance of evil-doers, led by Frieda, looks to take over Fairy Tale Land. But when Ella realizes her stepmother is out to ruin her storybook existence, she takes a dramatic turn and blossoms into the leader of the resistance effort.

In the 1400's, Rumpelstiltskin is imprisoned inside a small jade figurine. In modern-day Los Angeles, the recently widowed wife of a police officer, with baby in tow, finds her way into a witch's shop and purchases a certain figurine, resulting in the cackling beast being freed and demanding possession of the baby.