禁酒法時代のシカゴ。失業中のミュージシャン、ジェリーとジョーは偶然、殺人現場を目撃してしまい、ギャングのボス、スパッツから命を狙われるはめに。やむなく2人は女性だけの楽団に女装してもぐりこみ、寝台列車でマイアミに向かう。魅力的なウクレレ奏者シュガーにすっかり惚れ込んだ彼らだが、ジョーは、マイアミで富豪に変身して彼女に再接近を図る。一方のジェリーは、初老の富豪オズグッド三世から求愛を受けてしまう。

Heading east to Fort Worth to hire a schoolteacher for his frontier town home, Link Jones is stranded with singer Billie Ellis and gambler Sam Beasley when their train is held up. For shelter, Jones leads them to his nearby former home, where he was brought up an outlaw. Finding the gang still living in the shack, Jones pretends to be ready to return to a life crime.

Although she loves Christmas, Santa’s elf Christine (Summer Glau) wonders if there could be more to life than making toys in the North Pole. When Santa receives an emergency wish from a little boy in Los Angeles, he puts Christine on a special assignment, sending her to L.A. to give the VanCamp family—mom Sara (Eva La Rue), dad Scott (Dan Gauthier), and their two kids Ally (Izabela Vidovic) and Will (Mason Cook)—a holiday wake-up call. Will Christine fail her assignment and be sent back to the North Pole for good, or will the VanCamps come to their senses about the meaning of Christmas?