Leslie is disappointed when her new park plans are put on hold by a visit from two state auditors. April has plans for Andy at her birthday party, and Ann tries to piece together a romantic mystery.

When Ben prevents Leslie from holding a children’s concert, she takes matters into her own hands. Mark makes some big decisions about his life, and Andy gets some unexpected romantic attention.

Leslie and the Parks Department return to work after three months to discover that the budget crisis still looms. After unsuccessfully trying to have their budget restored by using Ann to charm Chris with a date, Leslie decides to make an all or nothing play by resurrecting the Pawnee Harvest Festival. Meanwhile, Ben restarts the youth basketball league and instates Ron and Andy to coach the only two teams. April returns from a vacation with a new boyfriend.

Leslie wants to bury a time capsule that encapsulates life in Pawnee, but an odd suggestion from a local man causes unforeseen consequences. Andy seeks help from Chris about how to win back April.

A local Pawnee group predicts the end of days, causing chaos in the Parks department.

Leslie and Ron briefly put aside their feud to help old foe Jeremy Jamm escape the clutches of the diabolical Tammy 2. Ben vows to help April discover her true passion in life, while Andy and Tom travel to Chicago.

Leslie and Ben suspect that Gryzzl is invading the privacy of Pawnee residents. April addresses the new Parks and Rec interns, while Tom helps Andy renegotiate a contract.

Leslie holds a rally in an attempt to keep JJ's Diner from closing. Tom surprises Donna with the ultimate pre-wedding gift.

Donna and Joe tie the knot, while April attempts to keep the wedding running smoothly. Ron creates an awkward situation between Tom and Lucy. Jen Barkley arrives at Ben and Leslie’s house with an interesting opportunity.

Ben and April conduct a series of interviews, while Andy and Donna help Ron adjust to a life change.

The Parks gang completes one last task together, before saying goodbye to Pawnee.