When Colleen teaches Matt her frugal ways, he gets hooked when he realizes how much saving money turns her on. Also, Tim attempts to fire Heather from her receptionist position at his office, John tries to break Sophia's fear of flying by taking her for a quick flight on a small plane, and Jen plans a quiet affair for Lark's first birthday, until the family takes over the celebration.

Dinner gets awkward for Jen and Matt when they find themselves alone without Greg and Colleen. Also, Lark accidentally locks out Greg and Jen from the house, prompting Greg to install a complicated home automation system; Heather and Tim try to sneak wine into an ice skating show to get out of paying at the venue; and John and Joan agree to an impromptu wedding after Colleen mentions that she's an ordained minister.

When Sophia asks Matt and Colleen about having children, an animated sequence featuring the whole family as farm animals reveals that they have been struggling to have a baby. Also, Jen and Greg's new neighbors aren't as nice as they initially seem; Heather interferes with Tim's hiring process at the office; and the whole family surprises Joan at her book reading, to her chagrin.

When Heather tells Sophia she has to get rid of her stuffed animals, Sophia stages a musical with them to see if she can change her mother's mind. Also, Colleen babysits Lark, and Jen comes home to find her dolled up in Colleen's old pageant dress; Joan's bike is stolen from the driveway so she enlists Matt, Greg and Tim to join her in tracking it down; and Tyler and Clementine get John's help to sell their homemade beef jerky at the farmer's market.