Betty struggles with a dilemma as she attempts to bail her family out of a financial crisis while juggling with the task of planning MODE's big assistants week at a medieval-themed restaurant...where Amanda's past ties to the establishment has a link and could come back to haunt her in a surprising way. On the romantic front involving the Meade siblings, Betty tries to give Daniel advice about his endless sexual addiction, while Alexis ponders about whether she should start taking up Rodrigo's offer. Meanwhile, Hilda prepares for her first exam to determine her future at cosmetology school as Wilhelmina prepares for her future plans to take over MODE as she makes even bolder moves for domination behind the scenes.

Betty gets a little overwhelmed about Wilhelmina's brand new hunky bodyguard. Plus, Henry returns to pay for insurance, which has Betty concerned about their relationship. Because of the fatal crash, Alexis gets amnesia and Bradford uses it to his advantage. Hilda comes to terms with recent events. And Amanda gets a surprise inheritance from her late biological mother.

Wilhelmina tries to becomes part of the Meade family by any means necessary. Amanda withholds evidence and Betty applies a deadly perfume made by Fey Sommers.

Betty finds herself in a dilemma between Henry and Gio when each makes a romantic offer to her. Meanwhile, a softball game between ELLE and Mode has rivals Wilhelmina and Naomi Campbell facing off in the field. Finally, Hilda makes quite a discovery about Coach Diaz, and a surprise visit from an orphaned French boy stirs things up at Mode.

Betty must make a life-altering decision about what to do with her life. Meanwhile, Ignacio lands a job at a fast food restaurant while Wilhelmina forces Daniel to make changes in his life.

A tragic accident leaves the Mode staff stunned and Betty must find out what really happened when a detective comes looking for answers. Meanwhile, Ignacio learns about Hilda and Coach Diaz.

Betty wants to be noticed by her cute neighbor, Jesse, by coming home later each night. Meanwhile, Amanda unexpectedly moves in with Betty after having nowhere else to go. Elsewhere, Marc and Cliff mull over the idea of moving in together.

Competition gets fierce when Betty and Marc both apply to YETI, the Young Editors Training Program that could be their ticket out of assistant-land.

Daniel and Wilhelmina make Mode's issue based on sex to spice things up. Matt is hesitant to take the next step with Betty. Marc and Amanda provide Betty with a makeover.

Just as Betty agrees to move in with Matt, Henry pays a visit to NYC, forcing Betty to admit she still has feelings for her former fiancé.

Betty finds that everything is changing as she begins her new job as associate features editor, especially since her boss is her ex-boyfriend, Matt Hartley, who is still unhappy about their breakup. In addition, Betty's new officemate, Megan, openly resents her, while Marc is constantly undermining her because he felt he was more deserving of the promotion. Meanwhile, without Betty as his assistant, Daniel is adrift, still mourning the loss of his wife, Molly. Betty tries to help Megan out on an assignment and connects with a thoughtful insect jewelry designer, Olivia, while Justin starts high school and is victimized by bullies-but won't tell his mom, Hilda. Claire Meade is finding that her power struggle at the company has her sitting pretty - for the moment -- opposite Wilhemina, who lost her claim to the Meade throne last season. But Wili has perhaps an even bigger problem than job security - she is harboring a secret at home that is driving Marc to distraction.

Betty gets to run her own Mode photo shoot at the United Nations to promote their anti-malaria initiative, "Nothing but Nets," but her big opportunity becomes a baptism by fire when Daniel's neediness takes its toll, and Matt openly demoralizes her. Meanwhile, Amanda does some soul-searching when she bonds with Daniel's new temp, Helen, a career receptionist as fabulous as Amanda is. Wili makes it her personal mission to protect a loved one, and Hilda reaches out to Marc to figure out how to help Justin.

Betty tries to get a scoop with Gucci's new designer, Evan York, through Hilda's customer, Sammy. Marc tries to stop her by getting Amanda and Matt to stage a showdown at Ignacio's restaurant. Daniel goes to a bereavement group and meets Natalie who shows him an interesting night. Wilhelmina flies to Bermuda to try and find Connor.

Wilhemina is livid when she lays eyes on drag queen impersonator - Wilhediva Hater. Betty and Daniel go through a rough patch as she wins an award for her blog. Daniel has the press hot on his heels. While Marc is pleased with himself when a famous director does a photo shoot with him.

The episode focuses on Betty Suarez struggling to tell her friend and boss, Daniel Meade about her new job in London. At the same time, Betty's sister, Hilda tries to keep their father from finding out she is moving to Manhattan. Wilhelmina Slater recovers from being shot by Tyler and Amanda Tanen finally finds her father.