It's the ZBZ Formal and Casey and Max decide to go together. At the event, Ashleigh and Rusty bond. Also, Rebecca decides to skip the event claiming she needs a break from sorority events.

Frannie and Casey sling mud at each other as the sorority presidential election approaches; Calvin faces the consequences after he cheats on Michael.

Casey faces a hard decision when she learns that Max is planning to go to Cal Tech for grad school and she is offered an internship. She turns to Lauren Conrad (who appears in a dream sequence) for advice. Also, the pledges are put through Hell Week and Rusty is driven to face the challenge no matter what he is forced to do.

Casey returns with secret plans to move out of the ZBZ house; a new freshman is highly desired as a pledge; Rusty tries to balance studying with life in the Kappa Tau house.

Now a sophomore, Rusty gets to participate in Rush Week for the first time and he is determined to land Andy as a pledge for Kappa Tau, but Calvin feels the pressure from Evan to get Andy to pledge Omega Chi. Who will prevail in the battle for the big man on campus? Ashleigh and Casey start to doubt Rebecca's "mole" abilities and begin to question whether she's actually a double-agent for Frannie's new sorority when the ZBZ girls’ rush week “Project Runway” fashion skit is successfully sabotaged by Frannie.

When Frannie challenges the sisterhood of ZBZ girls, and Casey retaliates by challenging the IKI’s to a game of dodgeball. Meanwhile, Rusty struggles with the fact that Dale is so in love with their new apartment that he never leaves, so Rusty enlists the Kappa Tau’s to invade the house to scare him away. But the plan backfires when the landlord shows up the next morning with a tour.

Rusty attempts to solve a mystery involving a secret service he ran across on campus in hopes of impressing Cappie. Meanwhile, some of Frannie's defectors from ZBZ house have asked to return, and Rebecca seeks dating advice from Calvin as she readies to go out with Robin.

After she falls into a manhole risking her life, coupled with the Kappa Tau's throwing an "End of the World" party, Casey is sent into an emotional tailspin, questioning her relationship with Max and her lingering feelings for Cappie. Meanwhile, Ashleigh faces her own punishment with Pan Hellenic after a complaint is lodged by Frannie for the Homecoming Float sabotage. Ashleigh turns to Casey to help fight the charge, but when Casey gets caught up in her own personal drama, she leaves Ashleigh to face Frannie alone. And with Dale having trouble keeping his cougar girlfriend's sexual advances at bay and Calvin trying his best to resist hooking up with his new hot roommate, Grant. Dale convinces Calvin to go in on a "purity pledge" partnership so they can help each resist their most recent temptations.

Casey finds out about Rebecca and Fischer's secret kiss, but when Rebecca confesses to Ashleigh, she outs Casey for keeping it a secret. When Mr. Cartwright, Mr. Howard, and Mr. Logan arrive for Father-Daughter weekend at ZBZ, they find out about their daughters' feud, and it reflects upon their weekend behavior. At the same event, Rusty gets nervous about meeting Jordan's father. Meanwhile, in an effort to get over their lost loves, Dale and Cappie fawn over the same girl, but they both ultimately get rejected.

It's Founders Weekend at the ZBZ House and Casey uses the opportunity to scrap for donations from the alums while Rusty tries to keep Jordan from dropping out of school.

In Casey and Ashleigh’s continuing struggle to bring ZBZ back to the top, Ashleigh decides a four-way mixer is the best way to gain their popularity back. Will her party save ZBZ from the fourth spot, or will she put them in even more debt?

Cappie and Casey go public, and Omega Chi and Kappa Tau get serious about pranking. Things heat up when Song Fest pits ZBZ against Gamma Psi. Rusty and Dale take their rivalry out of the lab.

The KTs and OXs are preparing for the Gods of Golf Tournament, the winner of which will be named the top fraternity on campus. Meanwhile, the ZBZs are disappointed to learn that the annual Greek Week has been canceled. Casey talks to Katherine, who agrees to bring back the Greek Goddesses Pageant, under the condition that the competition focus more on strength of character and less on looks, of course. Which houses will end the year on top?