Rebecca makes a surprise visit to Randall's home where she encounters his biological father, William. Kevin anxiously decides to take the plunge and move to New York to pursue a Broadway job, with the expectation that his sister, Kate, will move with him. Toby is determined to get Kate out of Kevin's shadow. Jack and Rebecca take home their babies from the hospital and adjust to the new life of parenthood, while coping with the loss of their other child. Rebecca struggles to connect with their adopted child and searches for answers.

Randall and William take a road trip to Memphis, where Randall learns about his biological father's past.

Deja's life with her mom, Shauna, and great grandmother Joyce unfolds from the very beginning. Randall and Beth welcome visitors to their home.

The Pearsons gather to celebrate their 40th birthday while musing about the day they were born.

Kevin and Madison navigate their engagement as Kate and Toby take a big step in their adoption journey. Jack and Rebecca brace for puberty.

Randall uncovers new truths about his past.

Kevin visits Randall in Philadelphia.