Lucy's situation grows desperate after she's taken captive by Flynn and brought to the 1893 Chicago World's Fair. In their dogged pursuit of Lucy, Wyatt and Rufus fall into a sinister trap.

Flynn teams up with legendary outlaw, Jesse James. Together they cut a bloody swath across the American frontier. Hot on Flynn's trail, Lucy, Wyatt, and Rufus enlist the help of an iconic Marshal to help bring the fugitives to justice.

Wyatt convinces Rufus to steal the Lifeboat for an unauthorized mission to prevent Wyatt's wife's killer from ever being born. Lucy must cover for them in the present, as Agent Christopher discovers the ship is missing and Anthony Bruhl makes an alarming confession.

As Lucy grapples with her newfound identity, Flynn disappears to 1927 France on the day that Charles Lindbergh completes his transatlantic flight. Now, Rufus, Lucy and their new soldier scramble through the City of Lights and team with young reporter, Ernest Hemingway, to keep history intact.

Flynn travels to 1931 Chicago to partner with gangster Al Capone. Lucy, Wyatt and Rufus, meanwhile, team up with the only man who can bring Capone down: federal law-enforcement agent Eliot Ness.

With one of our heroes' lives on the line, the team chases Flynn to 1954 through the paranoid corridors of Joe McCarthy's Washington D.C., en route to a secret, once-a-generation meeting of Rittenhouse leaders. While there, Lucy must wager the ultimate sacrifice to save history as we know it.