Go Eun-chan supports her family by working two jobs: as a Taekwondo instructor and a food delivery girl. While making a delivery, she meets Choi Han-gyul, the grandson of chairwoman Bang of Dongin Foods, a company that has a thriving coffee business. Han-gyul is a smart young man but he is fiercely independent and abhors the thought of being tied down by one career in his life. Sporting a short haircut, Eun-chan looks like a guy instead of a girl. Meanwhile, Chairwoman Park Soon-young arranges blind dates for her grandson Han-gyul to see him settle down.

Han-gyul still thinking she's a man, asks Eun-chan to pretend to be his "gay lover". She agrees to only show up on his blind dates, and he must buy her clothes and any time he touches her will be extra, and he has to give her an advance payment. Desperate, he agreed. Eun-chan has feelings for Han-gyul's cousin, Choi Han-sung, but then his ex-girlfriend Han Yoo-joo returns and says she wants to get back together with him. Han-gyul's grandmother owns a coffee shop and she forces him to visit it with here, and then she reveals his new assignment.

Han-gyul is forced to take over the Coffee Prince shop by his grandmother, and he enlists Eun-chan's help.

It is the grand opening for Coffee Prince, but business is harder to bring in than they expected.

The Coffee Princes head out to the countryside for their first team-building retreat, but one more person learns of Eun-chan's secret.

Eun-chan goes on her first "date" with Han-sung. Han-gyul struggles with his unexpected feelings for Eun-chan.

Han-gyul fires Eun-chan, which affects them, and all of the other Princes, more than either of them care to admit.

Still confused, Han-gyul proposes that he and Eun-chan become sworn brothers.

Han-gyul and Eun-chan celebrate their momentary bliss of brotherhood, but it is quickly shattered.

As the Coffee Prince Concert quickly approaches, the relationships between Eun-chan and Han-gyul, and Yoo-joo and Han-sung begin to fall apart.

Eun-chan wishes to reveal her true identity. Yoo-joo receives an offer from the States that threatens her relationship with Han-sung.

Betrayed on all sides, Han-gyul hesitates on forgiving Eun-chan. Meanwhile, Yoo-joo prepares to leave for New York.

Han-gyul and Eun-chan enjoy the "honeymoon phase" of their relationship, despite their awareness that he might leave for the States in a month.

Yoo-joo and Han-sung's wedding makes Han-gyul more anxious to marry Eun-chan.

Han-gyul's grandmother is in the hospital, but that doesn't stop her from meddling in his relationship with Eun-chan.

Eun-chan leaves for Italy to study to become a barista for two years.