Griff investigates its roots in Barmouth, discovers how the tiny cove of Mwnt copes with the impact of the modern world and considers the difficulties they face in deciding the fate of a medieval village on the Gower Peninsula which is in the process of being claimed by the sea.

In this programme, he visits Plas Newydd on the island of Anglesey in north west Wales, which the Trust acquired in the 1970s. Once it was the family home of the Marquis of Anglesey, and now property manager Nerys Jones has to think of new ways to attract visitors to this remote location. Griff investigates how this great stately home is run - and how it survives in these financially straitened times.

Surprisingly, the National Trust started in Wales and Griff Rhys Jones examines how it manages its farmland and tenanted farms across the country. Beginning in Pembrokeshire with the Trust's latest bequest, Treleddyd Fawr - the most painted and photographed cottage in Wales - Griff discusses the daunting restoration work facing National Trust building surveyor Nathan Goss. Through this derelict farm worker's cottage, as well as a working farm producing potatoes sold under National Trust branding and a farm which no longer farms but instead operates as a children's adventure camp, Griff explores the variety of ways in which the Trust approaches its guardianship of farms and farmland.