Jacks-of-all-trades Yun and Haberu investigate sounds emanating from an empty mansion. They locate the source nearby, while Mei hears the same tune.

Yun, Goro and their robot Jet Jaguar clash with a hostile pterosaur-like kaiju, Rodan. Mei travels to Tokyo to meet with a world-renowned researcher.

A legion of Rodan swarm towards Nigashio. The Otaki Factory crew scramble to lure them away from the city, but soon find themselves surrounded.

Yun and Haberu deduce that Misakioku may be linked to the Rodan incursion. Mei attends Li's lecture in Dubai, where the professor reveals new material.

Salunga emerges from the depths of the Upala research station shrouded in red dust. Meanwhile, Yun observes Anguirus exhibiting an uncanny ability.

Mei discovers a mysterious code in Professor Ashihara's research. The Otaki crew confront Anguirus as part of the city's extermination efforts.

Ground forces under the command of Matsubara respond to Godzilla's attack on Tokyo. Now in London, Mei attempts to decipher Ashihara's research.