Sawamura Matsugorou, the grandfather of a young Tsugaru shamisen player, Sawamura Setsu, has died. In losing the grandfather whom he'd always admired, Setsu has lost his own sound, and so he has gone to Tokyo with no plans other than to search for what he's lost. When he gets caught up in some trouble, he's saved by Tachiki Yuna, a woman who works at a hostess club. Events progress, and Setsu ends up unveiling his musical skills as the opening act for a live concert...

Setsu's unconventional mother, Sawamura Umeko, has suddenly descended and forced him to enroll at Umezono Academy. In helping to assemble a shamisen which has lain long disused, he meets Maeda Shuri, a girl in the same grade as him. While all this is going on, Setsu reunites with his older brother, who he hasn't seen in a while, and they play shamisen together...

After being so deeply moved by Setsu's performance, Shuri decides to seriously pursue playing the shamisen. With Setsu at the center, Shuri, Shuri's childhood friends Yamazato Yui and Yaguchi Kaito, and the addition of Nagamori Rai, a fellow student with shamisen experience in a different style, the Tsugaru Shamisen Appreciation Club is formed! But with everyone being new to Tsugaru shamisen, Setsu isn't sure how to approach teaching them, so he ends up...

The Tsugaru Shamisen Appreciation Club has entered the tournament that Umeko is sponsoring, the Matsugorou Cup. Also participating are Tanuma Mai, who has thought of Setsu as a rival since they were little, and her older brother, Tanuma Souichi. With strong competitors like Osaka's Kaji Takaomi and Fukuoka's Arakawa Ushio gathered from all over Japan, the tournament is shaping up to be a fierce battle. And now, the curtain rises on the Matsugorou Cup's group competition!

At last, it is time for Umezono Academy to perform. In spite of their nerves, the Tsugaru Shamisen Appreciation Club begins their performance boldly. Setsu's overwhelming sound overlaps and intermingles with the sounds of the other Umezono members who each hold their own. As the images that arise in their hearts come together, their sounds come together too. Where will their performance take them?

Kaji has been struck by a misfortunate accident—a string snapped in the middle of his performance. Seeing him continue on two strings to give an amazing performance stirs Setsu's heart. Then, Umeko reveals her true motives for creating the tournament to Setsu. Will Setsu play his grandfather's sound or his own sound? At last, his performance begins...

Setsu plays, following his heart. He sets an image within the notes. With all his heart, he tries to convey it. The sound that pours from Setsu' shamisen draws forth memories from everyone who hears it. Then, the favorite for first place, Tanuma Souichiro's performance begins. The sound is overwhelming. Waves of it shake the auditorium and his performance charms the audience. With both of their performances over, the conclusion of the tournament brings...