When Leslie decides to marry two penguins to promote the local zoo, she inadvertently causes an uproar when both penguins turn out to be male. Meanwhile, Mark puts Ann in an awkward situation when he asks her to go see a movie.

Leslie designs the Pawnee Winter Wonderland Festival, but has to hide from the press when a meeting with a disgraced Councilman lands her in the middle of a sex scandal. Meanwhile, Ron offers to cover for Leslie for the day, and quickly discovers how much work she really does.

After Jerry gets mugged by a bunch of kids, Leslie meets with the head park ranger in hopes of improving park safety. Meanwhile, everyone tries to be nicer to Jerry once he returns to the office.

Leslie hosts a late-night shift for a local telethon, and lands former Indiana Pacer Detlef Schrempf as a celebrity guest. Meanwhile, Mark has big plans for his relationship with Ann.

While advertising for the Harvest Festival, Ben has to deal with some troubling questions about his past. Meanwhile, Andy continues to try to win April back.

It's make it or break it for the Parks Department at the Harvest Festival which is full of surprises.

Leslie's book about Pawnee comes out just in time for her campaign. Ann tries to bond with Ron and April.

Leslie goes toe-to-toe with Bobby Newport in a debate. Ann, Tom, and Chris work in the media spin room, while Ron goes the extra mile to save a party for Leslie's donors.

Leslie helps a local swim team get more practice time by working on a new law that extends the hours at the community pool; Ron finds himself filling a pothole for a local woman (Lucy Lawless) as part of Chris' new plan to solve citizens' problems around the city; April and Ben take a road trip to Pawnee.

Leslie and Ben organize a black tie gala to raise money for their park, but their plans are derailed when a city-wide emergency response drill is called. Meanwhile, Andy prepares to take his Police Academy Exam and Ron fills in for Leslie on Pawnee Today.

Leslie's speech she prepared for a special event is stolen. Ben begins his new job at the Sweetums Charity Foundation. Ann summons the courage to ask an important question.

Leslie prepares a haunted house with Ben, Ann and Chris for Halloween on the day of the recall vote. Meanwhile, Tom fears the end of Rent-A-Swag and persuades Ron to help save his business.

Ben attempts to surprise Leslie with a special anniversary gift. Tom and Adam try to help Leslie bring positive light to the Pawnee-Eagleton merger, while April asserts her new authority as Donna’s boss.

Leslie aims to create a new town slogan, but runs afoul of local shock jocks. Andy scouts bands to play the Unity Concert and discovers an interesting secret about Ron, while Tom looks for a restaurant locale with Donna and April.

Leslie, Ben, and Andy head to San Francisco for a National Parks Conference. Tom opens his restaurant, Ron has another run-in with his ex-wife Tammy 2, Andy lives a rock and roll fantasy, and Leslie makes a huge decision about her future.

Leslie and Ron briefly put aside their feud to help old foe Jeremy Jamm escape the clutches of the diabolical Tammy 2. Ben vows to help April discover her true passion in life, while Andy and Tom travel to Chicago.

Leslie and Ben come under fire for participating in an old Pawnee tradition. April and Ron go on a scavenger hunt. Donna and Garry stroll down memory lane.

Ben and April conduct a series of interviews, while Andy and Donna help Ron adjust to a life change.