A grisly murder stokes class division on Snowpiercer, the Great Ark Train. Melanie Cavill, the powerful head of hospitality, deputizes a dangerous rebel to help solve the killing -- Andre Layton, the world's only surviving homicide detective.

The rebel forces are on their heels when a dangerous foe joins them. A plan is hatched to take the train, but it may destroy them all.

In the revolution's aftermath, Layton and Melanie realize the greatest threat to their survival is right over their shoulder.