In the aftermath of her recent assignment, Villanelle is ordered to take a break. Never one to do as she's told, she goes ahead with her next mission regardless. Meanwhile, Eve is given a dream opportunity to join a secret MI6 unit tracking Villanelle and the shady organization she works for.

Intel from Berlin points to the existence of a mole, prompting Eve to undertake her first surveillance operation. Meanwhile, Villanelle is sent to England to eliminate a member of British Intelligence. Villanelle wonders, could it be Eve?

Tensions are running high as the situation in Moscow escalates, and Eve starts to question who she can trust. Despite the danger, with her sights firmly set on Villanelle, Eve goes rogue. With both women under pressure, the stakes are higher than ever.

Villanelle's mission is proving anything but straightforward, forcing her to take drastic measures. Meanwhile, Eve makes a shocking discovery, which leads her right to Villanelle and an electrifying confrontation from which neither will emerge unscathed.

Villanelle's new circumstances are looking bleak, while Eve makes a dangerous deal with an old acquaintance. Villanelle is surprised when she is approached by a figure from her past.