Desna and her girls take pride in doing nails at the Nail Artisans of Manatee County, but amidst painting hooves and dreaming of a better life, she finds herself entangled in a money laundering scheme with the Dixie Mafia.
Haunted by Roller, Desna is desperate to get out from under Uncle Daddy's thumb. Bryce drifts deep into the Dixie Mafia. Polly reunites with a former gold-digging "friend," while Quiet Ann is in a lady-mess.
Desna and Jennifer may be Dr. Ken's only hope, as Bryce wrongly accuses him of a heinous crime. Virginia and Desna are forced to go to Polly for help to rid the world of two horrible birds from Desna's past, with one well-aimed stone.
All of our girls go to their new fancy salon for the first time. They aren't met with the kindness of strangers. Desna and crew decide to compete in Florida's premiere nail design competition for a big cash prize.
With a shocking revelation affecting the entire crew, Jennifer finds unexpected solace in her square-dancing instructor. Polly uses all of her powers of bad for good, for the first time, while Quiet Ann is torn about a big decision.
Desna goes searching for the truth with her crew right behind her, save for Jennifer, who is learning how to cope as a Dixie Mafia wife. When Desna's sleuthing hits pay dirt, she goes to the only person who can save her. Time is running out.
Following a harrowing escape from Roller's grasp, Desna is on the run. Roller is close behind, running from the Russians. In the end, only Desna can save Jennifer, Roller, and her crew with a sacrifice greater than any she's ever had to make.