When Nate thinks he has found a pattern to the anachronisms, it leads the Legends to London in 1897 to hunt down a time-traveling vampire. When they arrive in London, they run into Rip, but not everyone welcomes him back so quickly, leaving Sara to make a tough choice in the end. Amaya tries to connect with Zari, but she is still struggling with being part of the team. Meanwhile, Stein discovers what Ray and Jax are up to and is not entirely pleased.

A new fugitive is on the loose in 1951 Tokyo and Sara, Zari, Rory and Charlie try to capture it. As Constantine struggles to recover, Ray knows that magic might be the only thing that can save him but must look outside the team for help. Meanwhile, Ava joins Nate and his family for an unconventional Thanksgiving dinner.

When Mona discovers that her favorite author, Jane Austen, might be at the epicenter of a magical alert, Mona, Sara, Charlie and Zari find themselves in 1809. With Nate still dealing with family issues, Constantine and Rory perform a séance and receive a message from the other world that rocks Constantine to his core. Meanwhile, Ray and Nora are forced into close quarters.

When the Legends discover a new Encore, Sara, Rory and Ava end up crashing a high school reunion to attempt to stop a serial killer from finishing unfinished business. The Legends devise a plan to send Nate, Ray, Nora, Behrad and their hostage Zari, back in time to try and intervene in order to try to save everyone including them. Meanwhile, Constantine is annoyed that Charlie has been squatting in his old home, but it might be the blessing in disguise they were not looking for.

The Legends continue their hunt for pieces of the Loom of Fate and find themselves face to face with William Shakespeare. Ray, Nate, Rory, Constantine, Charlie and Behrad stay a little longer on the mission making things worse after lying to Sara. Meanwhile, Ray and Nora make another big life decision that leaves the team surprised and best buds, Nate and Ray, on the outs.

With Constantine obsessed with getting his powers back, he convinces Spooner to join him on the quest to find the Fountain of Imperium. After they find themselves at the end of the Spanish Civil War, Spooner makes an unexpected bond with someone she feels the need to protect. Meanwhile, Sara and Lita are concerned about Rory's health, so they try to convince him to take it seriously.

While the Legends contemplate how to get to New York City, Behrad discovers Hoover has train tickets to DC and Sara and Ava come up with a plan. With Zari stuck without the internet, she is forced to take a digital detox and becomes obsessed with figuring out who blew up the Waverider. Meanwhile, Astra and Spooner have stayed behind in Texas and are dealing with the aftermath of Astra's special spell.