When the Legends track down an anachronism in 1930s Hollywood, they discover it's none other than a time-displaced Helen of Troy and she's just started a war between two film studios. As the Legends try to fix history and return Helen to the Bronze Age, things get complicated when they are blindsided by the appearance of a former enemy. Sara contemplates an offer she is made, which would make the Legends leave the anachronisms be. Meanwhile, Stein and Jax find themselves in an unusual predicament.

When Sara decides to take some private time, the Legends pursue a lead on the whereabouts of the long-lost Earth Totem. Without Sara's knowledge, the Legends find themselves in 1717 to hunt for Blackbeard's long-lost treasure, but in true Legends fashion things go awry. Meanwhile, Ray feels guilty about something he has done and tries to make amends, which lands him in even bigger trouble.

A new fugitive is on the loose in 1951 Tokyo and Sara, Zari, Rory and Charlie try to capture it. As Constantine struggles to recover, Ray knows that magic might be the only thing that can save him but must look outside the team for help. Meanwhile, Ava joins Nate and his family for an unconventional Thanksgiving dinner.

When Hank commandeers the Waverider in 1973, the Legends find themselves on the run in an RV after kidnapping the President who is only able to tell the truth. With the help of Constantine, they discover the reason that none of them can lie making for an uncomfortable family road trip. Meanwhile, Nate and Zari, with the help of Nora and Gary, work together to find out what Hank is really up to.

With things getting out of control, Sara and Ava concoct a bold plan to take back the Time Bureau. But when things go awry, Zari and Charlie must work together to discover what Neron is planning. Elsewhere, Constantine is given a tough choice on who to save from Hell.

“Worlds lived, worlds died. Nothing will ever be the same.”

The crossover starts on Supergirl S05E09 Crisis on Infinite Earths: Part One (I), Batwoman S01E09 Crisis on Infinite Earths: Part Two (II), The Flash S06E09 Crisis on Infinite Earths: Part Three (III) and Arrow S08E08 Crisis on Infinite Earths: Part Four (IV)

The Legends find themselves going back to college and form a sorority in order to try to retrieve an ancient artifact called the Chalice of Dionyus that will help Charlie operate the Loom of Fate. Proud to be back at his college, Nate falls under Dion’s spell, letting him know of Sara, Astra, Ava and Charlie’s plan to get the cup. Meanwhile, on a father-daughter weekend, Rory takes Lita on a college tour, but Rory bails on Lita for an unexpected reason.