When Shelly's husband Ed goes missing, Sarah, Josh and Ali set out to find him, and instead find themselves lost in their past. Ali can no longer keep Maura's secret from Josh, which sends him into a tailspin. When Maura runs into a Pfefferman family friend she takes a stand for her new identity.

As they struggle to come to terms with Maura, Josh looks for answers from a rabbi named Raquel (Kathryn Hahn), and Ali looks for answers in a Gender Studies class with Syd. Sarah is left to answer questions from her own kids, who meet Maura for the first time.

Josh goes on a date with Rabbi Raquel while Ali explores her feminine side with a TA from Gender Studies class. The kids turn out for Maura's performance in a talent show, but her delight quickly turns to dejection.

Josh has a pool party for his new band, but he and his siblings are caught off-guard when Maura and Shelly arrive together; Ali reconnects with an old friend; Sarah finds her past catching up with her.

Josh and Raquel ask Colton to move in; Maura and Ali meet a feminist scholar—and Maura's former nemesis—at UCLA; Sarah launches a sexual fantasy life with a domineering figure from her past.

Davina and Shea inspire Maura to re-imagine her past; Colton's adopted family visits, leaving Josh facing a touch decision; Ali finds herself more drawn to Leslie; Sarah tries visiting a life coach.

Ali finds herself a little too close to Leslie when she seeks Leslie's advice on her UCLA application; Josh confronts Shelly about the past; Sarah uses a hard-won date night to explore some of her sexual fantasies.

Maura learns more about the trans experience; Josh heads down a reckless path; Ali and Sarah take Maura to the Idyllwild Wimmin's Music Festival.

Relationships are tested as the Pfefferman family gather for Maura's birthday. Sarah confronts Buzzy on temple board politics and Maura makes a big announcement, experiencing pushback from both Vicki and Shelly. The night culminates in a game of Sardines, forcing tense situations into tight spaces.

The Pfefferman family gathers at Hineni to support Sarah's alternative spiritual endeavor. After the news of a close friend's untimely passing, a distressed Josh leans on Raquel for support. Ali ends the night by witnessing the pitfalls of Leslie's prejudices.

Josh hits the road with an unlikely copilot, stirring up some undeniable chemistry along the way. Back at home, Shelly becomes wary of Buzzy's outlandish spending while Ali sees God during a nitrous induced trip at the dentist's office.

Maura and Ali uncover new truths about their family. Shelly gets bold in her improv class. Sparks fly between Sarah, Lila, and Len and they set off to hatch a plan. Josh is haunted by the past.

Maura revisits the night that Ali was born. Davina recalls memories of her youth and comes to terms with her troubled relationship with Sal. Davina and Maura each make a deal with God.

A dazzling two-hour movie musical fantasia. When the Pfeffermans face a life-changing loss, they begin a journey hilarious and melancholy, brazen and bold. As they face this new transition, they confront grief and come together to celebrate connection, joy, and transformation.