Rebekah arrives in New Orleans at her brother Elijah's insistence, but, concerned that Klaus is up to no good, she seeks help from a reluctant Sophie. Marcel enlists Davina's help.

In the war between Klaus and Marcel, Rebecca stands against her own brother Klaus. The chemistry between Elijah and Hayley gets stronger. Klaus accuses his siblings of distrust and betrayal.

Rebekah and Marcel have been trying to keep a secret from Klaus which may come to the light as we flashback to 1919. Elijah and Monique bond. Hayley takes Sabine hostage to try and get information on reversing the curse on her clan.

In order to show Elijah that her plan is what's best for him, Esther forces him to relive a time long ago when he loved a young woman named Tatia.

Rebekah becomes interested in a girl. Hayley struggles with coming clean and is surprised by something Jackson admits. Klaus learns something and heads to confront Hayley. Vincent sets his eyes on Marcel for answers to what Klaus is hiding.

Klaus must put aside his mistrust for Freya in order to save Rebekah's life.

With tension running as high as ever, Klaus and Elijah are forced to come together in order to uncover Lucien’s true motives in New Orleans. Cami is caught off guard when she finds herself face to face with Lucien and is surprised when he reveals some unexpected information about his past with Klaus. Elsewhere, a mysterious figure presents an enticing offer to Marcel, while Hayley turns to an unconventional way of coping with her current situation. Finally, an old acquaintance from Elijah’s past arrives in New Orleans and delivers some startling news involving the growing threat to the Mikaelson siblings.

In the aftermath of a horrific plan that left Cami dead, Klaus declares war on Aurora and Tristan. Elsewhere, Vincent, who has resumed his role as Regent to the New Orleans witches, is forced to use his magic against his will, while Hayley and Jackson ind themselves pawns in Tristan’s twisted game. Finally, after learning that Tristan is in possession of a powerful weapon that could take their family down once and for all, Klaus, Elijah and Freya enact a risky plan that leads to a tense showdown with The Strix.

When it's discovered that Freya has been kidnapped and is in the hands of a dangerous new threat, the Mikaelson brothers are forced to put aside their differences in order to save her before it's too late. Following a lead that brings them to Mystic Falls, Elijah and Finn run into deputy Matt Donovan, who does not take well to new vampires in his town. Meanwhile, Klaus and Hayley work together to uncover the motive behind one of Lucien's secret agendas, while a troubling shift in Kol's behavior leaves Davina concerned for their future together.

After five long years apart, Klaus attempts to reconnect with his daughter, Hope. Even as father and daughter bond, the rest of the family is still in turmoil. Elijah mediates a conflict between Hayley and Freya to determine the best way to protect the family. Meanwhile, in New Orleans, Vincent tells Marcel terrible secrets about his ex -wife, Eva Sinclair, even as they begin their search for a mysterious witch who plans to sacrifice a group of innocent children – including Hope Mikaelson.