Betty struggles with a dilemma as she attempts to bail her family out of a financial crisis while juggling with the task of planning MODE's big assistants week at a medieval-themed restaurant...where Amanda's past ties to the establishment has a link and could come back to haunt her in a surprising way. On the romantic front involving the Meade siblings, Betty tries to give Daniel advice about his endless sexual addiction, while Alexis ponders about whether she should start taking up Rodrigo's offer. Meanwhile, Hilda prepares for her first exam to determine her future at cosmetology school as Wilhelmina prepares for her future plans to take over MODE as she makes even bolder moves for domination behind the scenes.

Betty and Henry secretly go on a date to see Wicked, but Daniel, thinking that her date is Gio, gives two tickets to Betty and Gio and Gio does not want to give up those tickets. Meanwhile, Daniel gets in a meeting with an advertiser, a woman who has an interest in younger men, namely Daniel.

Betty is assigned to interview a novelist. Amanda and Marc may have uncovered who Amanda's biological father is. Daniel unknowingly dates Wilhelmina's sister.

Renee becomes convinced that Betty is in love with Daniel and thus starts a rivalry between the two for his affections. In the meantime, Gina returns to Queens to flash her new life and rich husband before Hilda's face.

A tragic accident leaves the Mode staff stunned and Betty must find out what really happened when a detective comes looking for answers. Meanwhile, Ignacio learns about Hilda and Coach Diaz.

Betty finds herself stuck between work and home life as she tries her best to appease both Hilda; Daniel, Molly, Connor and Wilhelmina.

Betty and Marc team up together to analyse Mode's budget. Matt continues with his pursuit to win Betty while she deals with some startling news on Facebook by a former flame

Betty finds that everything is changing as she begins her new job as associate features editor, especially since her boss is her ex-boyfriend, Matt Hartley, who is still unhappy about their breakup. In addition, Betty's new officemate, Megan, openly resents her, while Marc is constantly undermining her because he felt he was more deserving of the promotion. Meanwhile, without Betty as his assistant, Daniel is adrift, still mourning the loss of his wife, Molly. Betty tries to help Megan out on an assignment and connects with a thoughtful insect jewelry designer, Olivia, while Justin starts high school and is victimized by bullies-but won't tell his mom, Hilda. Claire Meade is finding that her power struggle at the company has her sitting pretty - for the moment -- opposite Wilhemina, who lost her claim to the Meade throne last season. But Wili has perhaps an even bigger problem than job security - she is harboring a secret at home that is driving Marc to distraction.

In the episode, Betty Suarez finally has her braces removed, but only after she enters a dream sequence in which she sees what her life would be like if she had had perfect teeth. At the same time, Justin Suarez tries to contact Austin, who he kissed in the previous episode and Daniel Meade secures the Million Dollar Bra for a MODE cover shoot. Meanwhile Amanda Tanen reveals that her relationship with Tyler is getting serious.