Daniel hatches up an idea to boost the Mode's financial situation after discovering that Betty and her family have been invited to the home of Matt's obscenely wealthy father, for an Easter egg hunt. Meanwhile, Wilhelmina discovers a reason to become suspicious of her son's DNA.

Baby William is the unlikely hold up in Mode receiving the money it needs to save the company, whilst Hilda is asked to help with a political campaign.

Just as Betty agrees to move in with Matt, Henry pays a visit to NYC, forcing Betty to admit she still has feelings for her former fiancé.

The sudden death of a Mode editor leaves Betty and Marc competing for the same job. Meanwhile, Matt lets his jealousy over Henry get the best of him.

Betty finds that everything is changing as she begins her new job as associate features editor, especially since her boss is her ex-boyfriend, Matt Hartley, who is still unhappy about their breakup. In addition, Betty's new officemate, Megan, openly resents her, while Marc is constantly undermining her because he felt he was more deserving of the promotion. Meanwhile, without Betty as his assistant, Daniel is adrift, still mourning the loss of his wife, Molly. Betty tries to help Megan out on an assignment and connects with a thoughtful insect jewelry designer, Olivia, while Justin starts high school and is victimized by bullies-but won't tell his mom, Hilda. Claire Meade is finding that her power struggle at the company has her sitting pretty - for the moment -- opposite Wilhemina, who lost her claim to the Meade throne last season. But Wili has perhaps an even bigger problem than job security - she is harboring a secret at home that is driving Marc to distraction.

Betty has Marc as be her temporary assistant thanks to Wilhelmina. Betty ends up covering the worst jobs in New York as an assignment but she needs to dress up as a hot dog and perform a Bollywood style dance. Marc has insecurities about how his idol at Vogue magazine will perceive him. Daniel is targeted by the leader of the Community.

At Wilhelmina's fund-raiser for Tibetan orphans Matt asks someone to be his partner. Daniel hires Natalie as his assistant, making Betty feel unneeded. Daniel finds out that Claire has a secret.

Ignacio prepares a romantic dinner for his Jean. Betty and Hilda take the Be-Shure instant pregnancy tests. Meanwhile, Hartley gets Denise Ludwig to fill in for Wilhemina. However Marc plans to bring Willie back. Claire with Amanda track down Tyler, the son she gave up for adoption.

When Betty becomes concerned that she and Matt are spending too much time together, she tries to give him something else to be passionate about -- a surprise gallery showing of his art work -- with unintended results. Meanwhile, Marc is horrified that his one-night stand, Troy, is falling for him, since Marc was Troy's "first", Hilda grapples with whether or not to tell Bobby that he's the father of her unborn child, and Cal officially replaces Wilhelmina with Denise.