In the 1600s, MacLeod sought refuge for a time in a monastery founded by Paul, another Immortal. There he encountered Kalas, an Immortal monk with a heavenly singing voice. When MacLeod discovered that Kalas was routinely taking the heads of Immortals as they left the sanctuary, MacLeod and Paul expelled Kalas from the monastery, separating him from the music that was his life. Now in the present, Paul and his choir have been lured out of their monastery for a concert tour. When Paul disappears after a concert, MacLeod discovers that Kalas is after his revenge. Meanwhile, two mysterious deaths at the hospital seem to be linked to negligence on Anne's part. Later, when drugs are found in Joe's bar, it becomes obvious that Kalas is trying to destroy MacLeod's friends before coming for him. MacLeod confronts Kalas and finds that Kalas is a strong and skillful fighter, better than MacLeod has faced before, and is nearly the victor. To save himself, MacLeod throws himself off the concert ha

MacLeod is picked up at the airport in France by his old friend, Hugh Fitzcairn. For the first time in the 350 years MacLeod has known him. Fitz has settled down as a Universitv Professor with the love of his life, Naomi. Flashbacks show MacLeod and Fitz's first meeting, when MacLeod was protecting the Doge's daughter from Fitz's amourous advances in Venice, 1637. When Naomi's jealous ex-lover is found strangled next to a computer displaying Fitz's falsified teaching credentials, Fitz goes on the run from the police. MacLeod realizes that Fitz is being framed by Kalas, who has followed him from the U.S. Kalas challenges Fitz and takes his head while MacLeod watches, unable to interfere.

When two Watchers end up dead by Kalas' hand, Joe realizes that Kalas is looking for Methos, the mythical ""oldest Immortal."" MacLeod knows that, with Methos' quickening, Kalas would finally be strong enough to defeat him. Kalas and MacLeod race to be the first to find Methos. Meanwhile, Richie muscles his way onto a top level motorcycle racing team and the champion, Basil, starts to get nervous. Flashbacks are to Paris in the 1920s, when Kalas, then known as Antonio Neri, was the toast of the opera world. When Kalas threatens a young girl in MacLeod's protection, they fight. Kalas escapes, but not before MacLeod inflicts a throat wound that destroys Kalas' vocal chords and the singing that has been his life since the Middle Ages. Kalas nearly takes Methos in battle and, realizing that he will not be able to defeat Kalas, Methos offers his own head to MacLeod. MacLeod refuses and challenges Kalas on his own, nearly defeating him when the police arrive to send Kalas to prison for the dea

Amanda accidentally helps Kalas escape from prison by trying to do MacLeod a favor and kill Kalas for him. After an attempt on Maurice's life thwarted by MacLeod, Kalas kidnaps Amanda to use as bait against MacLeod, but Amanda manages to escape. Meanwhile, Christine Salzer, the widow of a Watcher killed by Kalas (in ""Methos"") decides to get her revenge on Immortals and Watchers alike by exposing their secret to the media. Dawson and Methos team up to try and talk Christine out of it, but she's determined to take a computer disk with the identities of all known Immortals and Watchers on it to a newspaper publisher. Dawson, desperate, tries to kill her outside the newspaper building, but is stopped by MacLeod and Methos. Christine enters the building and the Immortals and the Watcher know their lives are about to change forever.

As Christine tells her tale to the newspaper publisher, MacLeod and Amanda, knowing their world is about to end, finally admit they love each other. Kalas kills Christine and the publisher and steals the computer disk before they have a chance to spread the story. Kalas offers MacLeod a deal -- MacLeod offers up his head or the contents of the disk are made public. Dawson and the Watchers attempt to find Kalas, but this only results in more dead Watchers. Methos tries to talk MacLeod out of it, but MacLeod agrees to fight Kalas on top of the Eiffel Tower. Kalas is defeated and the resulting Quickening, amplified by the Tower, sends a power surge that disrupts every computer in the vicinity -- including Kalas'. The information on the disk is destroyed and Immortals and Watchers maintain their anonymity.