While traveling through space in the Ark, the Autobots are ambushed by their enemies, the Decepticons. The battle moves into the Ark itself, leading to a damaging crash on Earth. After millions of years of dormancy, both sides are suddenly awakened and the battle on Earth is about to begin.

After being rescued from a Decepticon attack, Spike and Sparkplug offer their assistance to the Autobots. And the Autobots are going to need it in order to stop the Decepticons from gaining the power to conquer Cybertron.

The Autobots do whatever it takes to prevent the Decepticons from using a recently constructed ship to leave Earth and head for Cybertron.