During an especially bumpy flight Frasier tells a fellow passenger about his eventful past few weeks which are recounted in a long flashback. Bebe tells Frasier that he has been offered the chance to replace a recently deceased San Francisco radio host.

Daphne gives birth to a healthy son David. Frasier marries Martin and Ronee in the vet's office before Daphne and the baby leave for the hospital. Frasier grows very lonely with Martin out of the apartment and Daphne and Niles busy with their son.