Frasier hosts a party for the staff at the radio station to improve his image. Meanwhile, Roz dates a garbageman and Niles recalls his childhood memories with Daphne.

While Frasier and Niles uncover a human skull under the floor of their childhood home and let their imagination get the better of them, Roz's daughter's hamster has died and she is uncertain how to break it to her.

After Daphne and Niles elope in Reno, they are forced to go through several fake ceremonies in not to upset friends and family who wanted to be at the wedding.

Lilith comes to Seattle, and tells Frasier that she would like to have another child. She wants him to father the child through artificial insemination so that Frederick could have a full-sibling. Frasier is hesitant to go along with this, so Lilith tries to manipulate him by bringing up touching stories from Frederick's childhood.

Niles and Frasier plan a party to celebrate the unveiling of a painting that Niles has just purchased. However, Daphne notes that every event the brothers host inevitably ends in disaster.

KACL organizes a bike-a-thon to raise money for AIDS research. Frasier plans to just make a financial donation, but is goaded into participating after Julia announces her intention to take part. Daphne suggests that she and Niles also enter the event.

Roz tries to train the woefully incompetent Noel as Frasier's new producer as she prepares to leave KACL. She becomes increasingly melancholy about the possibility of leaving Frasier, and bristles upon learning of his new relationship with Julia.

Roz is disappointed when she is unable to get Alice tickets to a concert by children's performer Nanny G. After his family reveals that he used to be married to her, Frasier agrees to help Roz secure tickets.

During an especially bumpy flight Frasier tells a fellow passenger about his eventful past few weeks which are recounted in a long flashback. Bebe tells Frasier that he has been offered the chance to replace a recently deceased San Francisco radio host.