After losing Eri in a bet, Agri is forced to work for an old farmer in order to get her back from him.

The bizarre ability of a Warstar Insect makes Alata lose complete control of his invocation abilities, making his Gosei Cards become unpredictable!

Nozomu inquires the Goseiger about their current status after the fall of the Yuumajuu while a new enemy uses their research of the Goseiger to start their own invasion schemes.

Nozomu makes GoseiKnight question his purpose in order to convince him to help the Goseiger face down the Matrintis forces.

A time-controlling Matroid and a familiar opponent leads to a battle where Alata is thrown back to before the destruction of Heaven's Tower and the formation of the Goseiger team!

While the Goseigers deal with an onslaught from Robogog in the present, Alata must unite the three tribes to show their common bond to defeat Bakutofūji-ER in the past.

Brajira targets his former Headder Gosei Knight in order to force him to work for him again; while Agri and Moune must use their siblings abilities to defeat a threat that keeps healing itself.

Hyde takes offense at Brajira treating his Headders, including the now brainwashed Gosei Knight, as tools instead of as partners to work together with.

As Eri takes care of Nozomu, Alata and the other Goseiger try to handle Dark Gosei Knight, believing that he is the final key required for Brajira's Nega-End ceremony.