Claire decides to take on one of the villains herself, but she may not be as invulnerable as she believes. Nathan and Tracy search for the truth about their origins. Maya makes a shocking discovery. Hiro and Ando look for help in the most unlikely places.

Villains Knox and Daphne are on a quest to recruit Hiro and others to the Pinehearst fold. Angela and the Company are paralyzed from recent events. Peter goes to Pinehearst searching for answers but is stunned to learn who the founder is. Claire and Sandra go on a mission to save Meredith from the clutches of Doyle, a villain with abilities to control people mentally and physically. Nathan and Tracy try to trace their origins with Mohinder's help.

Trapped, Nathan and Tracy must fight to save their lives against Suresh, who has changed dramatically. Elle returns with shocking news for Claire and her family. Meanwhile, Angela tries to convince Sylar to save Peter; Daphne is ordered to kill Matt or she will be killed herself; and Hiro questions Usutu's advice.

Hiro attempts to delve into Arthur Petrelli's past to uncover the mystery behind him creating a rival company Pinehearst and his hatred towards Angela. Sylar stricken by guilt tries to stop his murderous ways with HRG and Elle vying for his blood. Meredith's training goes up in smoke with the arrival of her brother.