It has been several months since Alpha's reign of terror and Echo, who is now randomly picking up flashes of past engagements, weds a British businessman who has ties to one of Ballard's open FBI cases. Topher becomes the target of Dr. Saunder's aggression after she finds out that she is actually a doll named Whiskey. Boyd takes on a risky venture and mixes business with pleasure, while DeWitt makes Ballard an irresistible offer.

Echo is imprinted with the personality of a mother with a newborn baby. Complications arise when, thanks to Topher's programming, Echo takes too strongly to motherhood. Perrin ramps up his investigation into Rossum and DeWitt pays November a visit.

When Senator Perrin makes his stand against the Rossum Corporation, Echo attempts to stop his expose. Adelle, and Topher begin to see what they are up against in the rival of Washington DC.

Echo and Bennett have a shocking meeting as Adelle goes head-to-head with the ruthless head of the DC Dollhouse. Topher sees double when he must involve Victor in his espionage. Perrin finds a surprising witness to testify against the Rossum Corporation, but find he may be living in a house of cards.