Jess ærgrer sig over, at hun ikke kan tage sin klasse med til stranden, og lader det gå ud over en kinesisk restaurant, fordi dens menuer flyder i hele lejligheden.

Jess' vilde søster Abby kommer på besøg, og det ender katastrofalt. Nick har rollen som "wingman", da Schmidt snyder sig med til en bar mitzvah.

Schmidt hjælper Jess med at bruge en datingapp, og Nick, CeCe og Coach bliver skæve af marijuana-brownies, da Winston inviterer dem til en fest med andre politikadetter.

De sidder fast i lufthavnen på grund af flyforsinkelser, og vennerne er bange for, at de ikke når hjem til jul, men Jess får en særlig invitation.

Jess tager til en mindehøjtidelighed for en mand hun knapt nok kendte, Schmidt prøver at få sine bar mitzvah-penge fra sin mor, og Winston vil gerne være venner med Aly.

Jumping back into the dating pool, Jess finds herself dating a dud (guest star Taran Killam), but falls in love with his perfect parents (guest stars Henry Winkler and Julie Haggerty). Meanwhile, Nick (Jake Johnson) and Schmidt have very different ideas about how to run the bar.

Things heat up between Nick and Reagan over who can provide the loft with the best air conditioning unit. Meanwhile, Winston uses his “cop voice” to help Schmidt prep Cece for a newscaster audition.

Jess and the gang head to New York, where Schmidt believes he is being honored at his high school. Jess meets Det. Jake Peralta and helps him on a case, which leads her to Brooklyn’s 99th Precinct and a fateful encounter with Gina Linetti and Capt. Raymond Holt. Meanwhile, Nick and Winston scramble to make subway fare, while trying to impress Det. Charles Boyle, and Cece faces off with her new mother-in-law.

The crossover starts on Brooklyn Nine-Nine S04E04 The Night Shift (I).