Inspired the by the students, Will composes a mash-up for a very special occasion. Meanwhile, in a shocking reversal of roles, McKinley High’s cool kids are dethroned in an “icy” fashion, and Sue shows off her softer side.

When the Glee Club is left out of the yearbook because of budget cuts, everyone seems to be relieved that they won’t have to suffer the ridicule of years past. Everyone except Rachel, that is, who wants her legacy cemented in print. Meanwhile, Quinn makes it her mission to be in the Cheerios yearbook photo, and Will and Terri’s relationship is shaken to its core.

Fresh off their monumental win at Sectionals, New Directions is on top of the world. With an extra spring in their step, they get to work on new numbers for regionals. But when Rachel and Finn’s budding romance hits a bump in the road after she becomes involved with a competitor, Jesse St. James, it threatens to derail the entire club. Mr. Schuester pays a visit to Shelby Corcoran, coach of Vocal Adrenaline, to confirm Jesse’s motives. Meanwhile, Will is newly single, but is he ready to take his relationship with Emma to the next level? And to top it all off, Sue Sylvester returns from her condo in Boca with a renewed focus on destroying Will and disbanding the glee club.

Mr. Schuester starts an investigation when a salacious list about members of the glee club circulates the halls of McKinley High. Meanwhile, Sue is mortified when a video of her performing the Olivia Newton-John classic "Physical" surfaces

Mr. Schuester encourages the glee club to express themselves using the music of Lady Gaga. Meanwhile, Rachael has a life-changing encounter, and Tina embraces her inner Goth.

After a tough loss at Regionals, the glee club is back as the underdogs of McKinley High. When the school is faced with even more budget cuts, Mr. Schuester pushes the kids to help recruit new members for the club. Rachel and Finn disagree on new recruits, and aren’t sure they will be able to come together in perfect harmony.

When Brittany and the gang try to convince Mr. Schuester to let them do a Britney Spears number, he is reluctant to go that edgy. Meanwhile, an insecure Will is curious about Emma’s new beau, Dr. Carl Howell, so he convinces her to bring him in to talk to the kids about dental hygiene. While the dreamy doc inspires the girls to take their dental health more seriously, Will is left feeling worse than before.

The glee club works on a duet assignment, and Finn and Rachel plan to throw the competition so that Sam can have a shot at winning. Meanwhile, a love triangle between Mike, Tina and Artie develops.

Will decides to have the glee club perform “The Rocky Horror Show” for the annual school musical after learning of Emma’s newfound love for the cult classic. Despite Sue’s usual scheming, the members of New Directions inject their distinctive style to such timeless tracks as “The Time Warp” and “Damn It, Janet.”

After Finn and Sam find an unconventional way of controlling their urges, a few glee club members pick up on the strategy and start using it too. But when Coach Beiste gets wind of what they are up to, the glee club has some explaining to do. Meanwhile, the Girls vs. Boys Tournament returns, Kurt meets an ally from a rival glee club, Puck and Artie join forces and coach Beast is on the verge of quitting her job.

When Mr. Schuester gets sick, a substitute teacher takes over his Spanish class and the glee club and winds up winning over the whole New Directions clan. Meanwhile, when Sue fills in for Principal Figgins as McKinley’s top dog, she tries to flex her power on campus.

It's all about wedding and parents with Kurt getting ready for his father Burt's upcoming marriage to Finn's mom Carole, Sue decides to compete by getting married as well which will lead to the introduction of an awesome guest star: Carol Burnett. Carol will play Sue's Nazi-hunting mother Doris, who comes to visit Sue in Ohio.

New Directions heads to the big Sectionals competition to go head-to-head against the Warblers and the Hipsters. While trouble brews between some of the young couples, Emma and Carl grow closer.

It's Christmas time at McKinley High and Finn tries to get everyone in the holiday spirit; Brittany needs no help because she still believes in Santa Claus. In other news, Sue scams the teacher's Secret Santa

A local football championship game takes place and the "Thriller" mash-up is performed during the halftime show. The Cheerios, McKinley High's cheerleading team, compete at Regionals.

The glee club agrees to help raise funds for another under-the-radar group; big decisions are made and relationships are tested.

The glee club learns about self-acceptance through the music of Lady Gaga; Lauren and Quinn compete for prom queen; Ms. Pillsbury faces with her issues; Rachel makes a life-changing decision; Santana schemes.

When the entertainment for prom falls through, Principal Figgins asks New Directions to perform at the dance much to prom coordinator Sue’s dismay. Things get heated as the competition for prom king and queen draws closer, and Jessie St. James returns.

The school musical West Side Story is readied for performance, with Artie as lead director. Finn meets with a football recruiter, whom Coach Beiste gains as an admirer. Kurt and Blaine reach a new, physical stage in their relationship, as do Rachel and Finn in theirs.

In honor of Valentine's Day, Will challenges the New Directions to find and perform the world's greatest love songs. Meanwhile, Rachel's two dads Hiram and LeRoy Berry pay a visit to McKinley High.

Things heat up as the New Directions go head-to-head with the Dalton Academy Warblers at Regionals.

The gang must rally an uncharacteristically insecure Rachel in time to make her Broadway debut in Funny Girl… but not everyone in the audience is wishing Rachel the best.

After Rachel's humiliating failure as TV actress, she comes home to Lima to figure out what she wants to do next. Upon discovering that Sue has banished the arts at McKinley, Rachel takes it upon herself to reinstate and lead the Glee Club. Meanwhile, Blaine, no longer in a relationship with Kurt, has moved home to coach the Warblers while Will is coaching rival Vocal Adrenaline, and Sam is the assistant coach for the McKinley football team.

Puck, Quinn, Santana, Mercedes, Brittany, Artie and Tina all return to McKinley for Homecoming and help Rachel and Kurt recruit for the new glee club.

Rachel and Kurt struggle as they try to rebuild and co-direct the glee club at McKinley. Meanwhile, Sue is concerned when Becky brings her new boyfriend to town.

Rachel hopes to form a deal in which Will's glee club purposefully under-performs at their invitational; Sue wants Sam to distract Rachel.

A new job forces Will to take a hard look at the choices he’s made. Rachel must come to terms with the fact her family home has finally been sold. Coach Beiste, returns to McKinley and the romantic tension that has been brewing between Sam and Rachel finally boils over.

GLEE goes back in time to see why the original members of New Directions signed up for glee club in the first place.

All of Will’s hard work and dedication to the Glee Club and to fostering the talents and dreams of his students pays off in a major way, and everyone shares in the feeling of joy that permeates the school as teachers and students reflect on their shared journey. Old rifts and wounds are healed and new sights are set on a promising future. Everyone seems to be ending up exactly as expected and right where they belong… along with more than a few unexpected surprises.