When Richard Mayhew helps a wounded girl on the street, he finds himself beginning a nightmare journey that plunges him into a weird parallel world underneath London. The girl is on the run from two cut-throats who she thinks killed her family. But why are they after her? And now that Richard is involved, will he ever return to a normal life?

Angel Islington is the only person who can tell Door who killed her family and help get Richard back to his old life. But how do they get to him? The Marquis suggests a visit to Earl's Court - but it has a worrying tendency to move around. Meanwhile, Croup and Vandemar alert Door to the fact that there is a traitor in her midst... but who?

The Angel Islington has charged Richard and Door with finding a sacred key that will answer all their questions. But getting it involves the dinner party from hell, and an ordeal from somewhere worse. Meanwhile, Croup and Vandemar show their tastes for the finer things in life while developing an unhealthy interest in the Marquis.

With the sacred key in his possession, only one person stands in the way between the Angel and his ultimate ambition - Door. As the apocalyptic climax looms, everyone faces their own destinies.