When a scandalous story in the college newspaper hits stands trashing the Greek system, the brothers and sisters of Omega Chia, Zeta Beta Zeta and even Kappa Tau are in a frenzy to figure out who the undercover culprit could be. Meanwhile, after navigating some troubled waters with her boyfriend, Ashleigh turns to Calvin for support and even goes so far as to hit on him, only to learn he has a secret.

Rusty and Casey return to CRU but must contend with the backlash from Jen K's story, and deal with the fallout from their relationships. Meanwhile, Cappie and Rebecca must make a choice about their relationship.

Kappa Tau and Omega Chi face off in an epic battle of practical jokes complete with waterguns, duct tape and farm animals. Rusty tries to mediate the situation to save his friendship with Calvin but finds it only escalates the rivalry. Meanwhile, Casey learns that the only way to rid ZBZ of Lizzi, the conservative national rep overseeing them, is to forgive Frannie and reinstate her in a "diamond" ceremony.

Trouble is caused by Casey when she bends the sorority house rules for Ashleigh. Cappie tries to get rid of restrictions.

Meanwhile, Rusty finds himself attracted to Tina, the girl who wants to take down the Greek system.

Omega Chi and Kappa Tau each throw parties to celebrate the end of the campus restrictions.

The students enjoy spring break on the beach. Cappie and Casey explore their new friendship at a time when Rebecca needs Cappie's attention. Evan looks to move on and escape his feelings and ends up with a familiar person. Rusty returns to school and on his way back he reunites with old friends.

Season 2 begins with the annual Greek Week competition. Despite their opposing houses, Calvin and Rusty strive to remain friends. Meanwhile, Casey has to decide what to do about Rebecca's actions over spring break, and Rebecca's continuous bad behavior doesn't do the situation any good.