Shocked to learn that ZBZ has slipped in the ranks down to fourth, Casey is determined to bring ZBZ back to the number one spot by motivating their pledge class to take action, believing that, if they create a cool photo shoot for the Lamda Sig calendar, it could improve their house’s ranking. However new pldge Abby convinces the other pledges to boycott the shoot so, in order to make nice with the ZBZ pledges, Ashley and Casey get them all makeovers. Meanwhile, dissatisfied with her college experience, Jordan wonders how to make her time at CRU more fulfilling and, in an effort to prove his love and commitment for Jordan Rusty decides to lavaliere her. When Rusty divulges his plans to his KT brothers, they warn him of the Kappa Tau curse that occurs when a brother lavalieres his girlfriend. Elsewhere, Cappie helps Evan find a part time job to help his financial situation, but when he meets one of Evan’s new and very attractive co-workers, Lana, he decides to get a job too.

In Casey and Ashleigh’s continuing struggle to bring ZBZ back to the top, Ashleigh decides a four-way mixer is the best way to gain their popularity back. Will her party save ZBZ from the fourth spot, or will she put them in even more debt?

Cappie and Casey go public, and Omega Chi and Kappa Tau get serious about pranking. Things heat up when Song Fest pits ZBZ against Gamma Psi. Rusty and Dale take their rivalry out of the lab.

Casey is ready to discuss the future with a resistant Cappie, and the situation isn't helped by handsome Washington politician Joel (Sam Page), who encourages Casey to envision a potential career in politics after she volunteers to help him with his voter registration campaign at the Cyprus Rhodes campus. Find out if Cappie is ready to grow up or if he will fall behind.

The KTs and OXs are preparing for the Gods of Golf Tournament, the winner of which will be named the top fraternity on campus. Meanwhile, the ZBZs are disappointed to learn that the annual Greek Week has been canceled. Casey talks to Katherine, who agrees to bring back the Greek Goddesses Pageant, under the condition that the competition focus more on strength of character and less on looks, of course. Which houses will end the year on top?

The students of CRU are on Spring Break in Myrtle Beach, which means the year is winding down and the seniors have big decisions looming over their heads. As Ashleigh contemplates her job options, Casey is on edge waiting to see where she'll be accepted to law school. While Cappie is busy planning a romantic surprise for Casey's birthday, she worries that not making it into CRU Law could mean the demise of their relationship. Will Cappie and Casey survive Spring Break intact?

The season opens with Casey, Evan, & Ashleigh’s graduation day. Casey and Cappie haven’t spoken since Spring Break and are anticipating saying their final goodbyes to each other, which does not go as smoothly as expected. We then skip to the end of summer and Casey packing up to move to Washington DC for law school. Through some prompting from Evan, Casey learns that the reason she did not get into CRU law school was a result of a bad recommendation from Joel (Sam Page). After Casey confronts Sam, he reconsiders and has his letter rescinded. Casey is accepted into CRU law and decides to stay after all. Ashleigh has moved to New York for an internship that is not exactly what she was hoping for. Evan is attending CRU law school with Casey.

The KT’s decide since they lost several members of their frat last year, it’s time to actually try to recruit pledges. The KTs set up a booth during rush on campus to help with recruiting. Tegan (Charisma Carpenter) shows up to help the ZBZs maintain their newly regained best house on campus title. Tegan and Rebecca argue on how rushing should go and Casey comes to Rebecca’s rescue with a plan to have Dana (Martha MacIsaac) keep Tegan distracted. The ZBZs entertain a potential new pledge Heather, who is Heath’s sister. Dale also entertains the idea of rushing this semester

Rusty is planning Calvin‘s 21st birthday party at his apartment. Casey and Ashleigh have not been speaking and are afraid to run into each other at the party. A really bad snowstorm hits Cyprus Rhodes shutting down all roads. Since Heath is not able to make it back with the alcohol, the gang of eight decides to walk to Dobbler‘s. Inside Dobbler‘s, the group decides to play the game "Kiss & Tell" where secrets and feelings are revealed.

In the series finale, the Kappa Taus try to stop the demolition of their house. Meanwhile, Cappie and Casey come to a realization about their on-again, off-again romance; and Cappie and Evan determine whether their friendship is meant to last.