Shunsuke Aoba has just transferred to a new school and can’t wait to attend. Excitedly, he sprints to the school but, upon arrival, finds that no one is there. Unfortunately, Shunsuke didn’t know that his first day was a holiday, of all things. However, it seems that he is not the only one on campus. As he is about to eat his lunch, he meets a student named Natsumi. Shunsuke gives her some of his food, and the two strike up a conversation. Natsumi was born as a so-called SL girl, a particularly tall girl. She herself accepts her specialness but is rejected by many of her classmates. How will Shunsuke react to this? Will their initial friendship continue?

Tanjiro Kamado vive feliz con su madre y hermanos pequeños en la montaña, subsistiendo humildemente. Una mañana, al regresar a casa, se encuentra a toda su familia asesinada y a su hermana Nezuko convertida en demonio. Tanjiro buscará desesperadamente la manera de devolverla a su condición humana y de cobrarse venganza. Para ello, se someterá a un arduo entrenamiento, con el objetivo de ingresar en el Cuerpo de Matademonios y dar caza al demonio Muzan Kibutsuji, con la ayuda de Nezuko, que se resiste a perder del todo su humanidad. Juntos forman la nueva hornada de los guardianes de la noche.